C. Hart Merriam
Clinton Hart Merriam was born in upstate New York on 5 December
1855. His family was a wealthy, successful family with a large estate
surrounded by forests brimming with a variety of plants and animals.
This atmosphere allowed Merriam to spend many hours in the woods,
exploring the local flora and fauna. This early exposure encouraged
Merriam to have a deep appreciation for nature.
Merriam was educated at Yale University and studied human anatomy.
This education culminated in completing medical school and becoming
a physician. Many early naturalists were in the medical profession,
and Merriam used this time to go on field studies and collect vast
amounts of biological information. At the age of 18, he published
his first summaries of his biological studies.
In 1886, Merriam began work for the United States Department of
Agriculture in the Entomology Division. He used this platform to
promote in-depth biological surveys of the United States. Merriam
felt this would discover new organisms in America and eventually
lead to biogeographical studies, investigating interactions between
biota and abiotic elements such as temperature and elevation.
Merriam first visited the Colorado Plateau in 1889 while on the
Arizona Territory San Francisco Mountain Region Survey for the USDA.
He chose this region to investigate biogeographical interactions
because "different climates and zones of animal and vegetable
life succeed each other from base to summit" and "because
of its southern position, isolation, great altitude, and proximity
to an arid desert." Merriam trekked greatly around and on the
San Francisco Peaks, including the cinder hills, Walnut Canyon, Grand
Canyon, and the Painted Desert. His expeditions included extensive
plant and animal collections and elevation documentation from an
aneroid barometer. These studies eventually supported his Life Zones
concept of biogeographical elevation gradients.
Life Zones Concept as Proposed by Merriam
|
| Warm Desert |
< 1500 m |
High temperature, low precipitation desert with cacti and
desert flora and fauna. |
Desert Grassland /
Pinyon-Juniper Woodlands |
1500 - 2000 m |
Extremes of temperature and precipitation. Open landscape with Colorado
Pinyon Pine and one-seed juniper trees. Pinyon jay and large rock squirrel
common to the area. |
| Ponderosa Pine Forest |
2000 - 2500 m |
Largest stand of ponderosa pine trees in North America. Seasons characterized
into long winters, harsh droughts, and strong monsoons. Fire plays an extensive
role in forest ecology. Abert Squirrel, Rocky Mountain Elk, Steller's Jay,
and broad tailed hummingbirds found extensively in this region. |
| Mixed Conifer Forest |
2500 - 3000 m |
Rich moist soil supports stands of Douglas and white fire with limber
pine and quaking aspen. Black bears, mule deers, and elk can be found in
this zone. |
| Spruce-fir Forest |
3000 - 3500 m |
Heavy shade and high precipitation supports mainly Engelmann Spruce and
corkbark fir with extensive high-altitude meadows. Red squirrels, gray-collared
chipmunks, Clark's nutcrackers, and red-faced warblers are found in this
zone. |
| Sub-alpine |
~ 3500 m |
At the timberline, dwarf Engelmann spruce and bristlecone pine cling
to the mountain in a harsh environment. The water pipit can be found at
the timberline and bighorn sheep used to be found there as well. |
| Alpine Tundra |
3500 - 3852 m |
Extreme cold and high winds make this a harsh, brutal environment with
small flowering plants and few animals. The San Francisco Peaks groundsel
is found only in this zone and is currently under threatened status. |
|